Beta diversity measures
The following beta diversity measures are available:
- Bray-Curtis:
- Jaccard:
- Euclidean:
where

is the number of OTUs and

and

are the abundances of OTU

in samples

and

, respectively.
The following distances are also available:
- Unweighted UniFrac:
- Weighted UniFrac:
- Weighted UniFrac not normalized:
- D_0 UniFrac: The generalized UniFrac distance
- D_0.5 UniFrac: The generalized UniFrac distance
where

is the number of branches in the phylogenetic tree,

is the length of branch

;

and

are the proportion of taxa descending from branch

for samples

and

, respectively; and the indicator functions

and

assume the value of

if any taxa descending from branch

is present in samples

and

, respectively, or
0 otherwise.